More specifically, it requires the completion of the Sanctum of Domination achievement for the teleporter option to show up after all required steps have been unlocked for it.
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Sanctum of Domination Clear Required for Desmotaeron Teleporter
Blizzard has published a support article about the missing Desmotaeron animaflow teleporter, revealing that the teleport is only available to characters that have killed all 10 bosses in Sanctum of Domination.
